The opening item in C.P.E. Bach's most original group of symphonies begins with an Allegro di molto taking off from a single, stabbing note repeated several times, increasing in speed, density, and loudness into a full theme, then dropping suddenly to piano, and rushing back in full force. These patterns repeat with great variety and ingenuity throughout the movement. This is a prime example of Sturm und Drang writing outside the works of Haydn, and includes a few woodwind passages that look ahead to the delicacy of Mozart. The movement ends with a slow section that leads smoothly into the Largo, a miniature by comparison. This is music of great patience and sensitivity, coupling the flutes with the lowest string instruments for an unusual sonority. The movement breaks off almost in mid-phrase, whereupon the finale, Presto, bursts in. Fast and ebullient, the movement is based on a flamboyant opening gesture and rapid string passagework that in other hands might seem mechanical. C.P.E. Bach's quick shifts of mood and sudden bursts of humor keep the music engaging, despite its modest basic materials.

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